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When Selective Trout was first published in 1971, Swisher and Richards included Drunella lata (Small Blue-Winged Olive, Slate-Winged Olive) as a Midwestern "superhatch." Although it can also be found in many Eastern trout streams, it is probably more important to Midwestern anglers. Typically a morning emerger, this species often competes for the attention of trout with more abundant Tricorythodes and small baetids during parts of July and August. For this reason, the authors of Selective Trout considered the concentrated evening spinner falls to be more important than the somewhat sporadic morning emergence. From an angling standpoint, this situation is nearly the opposite of the earlier Drunella cornuta emergence in the East, where the morning emergence is usually the main event and spinner falls are often of little consequence.
Currently, Drunella lata shares its name with another mayfly, the former D. longicornis. That mayfly can be important in mountainous areas in the Southeast, but they are larger and the nymphs lack the distinctive pale markings mentioned in the Juvenile Characteristics section. (The information on this page does not describe D. longicornis) Read more...
DarkDun | March 4th, 2007, 9:40 am | |
Posts: 16 | The D.Lata emerges in Michigan waters at 10 AM on the dot and stops at noon from about June 25 thru July 10. I have fished this hatch avidly for years and find it very punctual on moderately overcast days. Sunny days make it much shorter duration, about 30 minutes. A size 14 imitates it perfectly with dark dun wings, bright olive green body and med dun tails and legs at emergence. The body color does change to dark green after a while. The D.Lata also is significant in PA Northern Streams in Mid May. I have not encountered it in the South Appallacian streams as yet. I fish some smaller BWO (#16-20) in NC but none so large as D. Lata. | |
Troutnut | March 4th, 2007, 10:42 am | |
Administrator Bellevue, WAPosts: 2737 | Thanks for sharing! As I wrote in the description, I think local advice like yours is even more important for this species than for most others. | |
Jason Neuswanger, Ph.D. Troutnut and salmonid ecologist | ||
Konchu | March 7th, 2007, 4:52 am | |
Site Editor IndianaPosts: 505 | How easy is it to recognize the hatches of the old species combined under Lata? Are these mainly local differences? I've never been able to sort them out very well at any one place. | |
Taxon | March 7th, 2007, 11:21 am | |
Site Editor Royse City, TXPosts: 1350 | Konchu- The former Drunella species now considered to all be Drunella lata are: D. cornuta, D. cornutella, D. longicornis, and D. lata. As mature nymphs, and under magnification, their frontal horn differences are quite distinctive. As duns and spinners, they would be extremely difficult to reliably differentiate. | |
Best regards, Roger Rohrbeck www.FlyfishingEntomology.com | ||
Konchu | March 7th, 2007, 8:08 pm | |
Site Editor IndianaPosts: 505 | My question was more about the behavior, emergence time, color, ANYTHING about the spinners and duns. | |
Taxon | March 7th, 2007, 10:55 pm | |
Site Editor Royse City, TXPosts: 1350 | Konchu- Okay, Konchu, I’ll take a crack at answering your question (as clarified), but please understand, Drunella lata doesn’t reside in the Pacific NW, so I have no experience with them, and am strictly relying on the considerable book knowledge to which I’ve been exposed. Hopefully, Jason, Gonzo, Louis, and others can provide some knowledge based on first hand experience. Anyway, with regard to the Drunella lata (subordinate species name equivalent) duns, Knopp/Cormier lists their dates of emergence as follows: D. cornuta – 1st week of May through 2nd week of July D. longicornis – 1st week of June through 2nd week of July D. lata – last week of June to mid August D. cornutella – 1st week of July through 1st week of August The time of emergence starts around noon in early May, and as the season progresses, and morning temperatures rise, starts at 5 AM by mid July, at which point it reverses itself, and starts progressively later in the morning, 8 AM by early September. In preparation for emergence, the mature nymphs are said to “excitedly” crawl to slower water, often loosing their footing and becoming vulnerable to feeding trout as they drift helplessly with the current. Trout are said to show consistent preference to emergers over duns, both actuals and imitations. D. cornutella and D. longicornis duns are colored light olive green when emerging, and medium olive when aged, have gray to dark slate wings, and legs with ambur with an olive cast. D. lata duns are colored light olive when emerging, and medium olive when aged, have dark gray wings, and legs are ginger with an olive cast. D. cornutella duns are colored chartreuse when emerging, quickly turn to light olive, medium olive when aged, have bluish gray wings, and legs are olive. With regard to spinners, mating swarms form an hour before dusk, and female spinners return to the water for egg laying, sometimes resulting in impressive spinner falls (for about a half hour before dark). The spinner falls are said to be of minimal interest to the trout, other than in mid July, when other mayflies are absent. | |
Best regards, Roger Rohrbeck www.FlyfishingEntomology.com | ||
Martinlf | March 8th, 2007, 5:44 am | |
Moderator Palmyra PAPosts: 3233 | I only have experience with Cornuta locally here in PA. Roger's general observations fit my experiences with one exception. On sunny days I have seen Cornutas emerge from about 8:30 or 9 until around 10, though on drizzly days I've watched them emerge almost all day long. I have noted hatches from May 25 to June 3. I believe that hatch times vary by location, and time of year, though. Gonzo knows a lot more about Cornutas than I do, and can shed more light on other Lata emergences also. | |
"He spread them a yard and a half. 'And every one that got away is this big.'" --Fred Chappell | ||
Konchu | March 8th, 2007, 10:26 am | |
Site Editor IndianaPosts: 505 | Sometimes I've seen hatch DATES vary with altitude in eastern Tennessee and parts of North Carolina. | |
Troutnut | March 8th, 2007, 2:59 pm | |
Administrator Bellevue, WAPosts: 2737 | Konchu that's a big can of worms you opened. :) I have no idea to what extent the behavioral variability within the species is actually associated with the former species boundaries. However, I do have LOTS of notes from anglers' books about them. This will give everyone a little peek into how I wrote all the articles about all the hatches last summer. I read and highlighted relevant passages from 8 different angler-entomology books, then I typed ALL of the highlighted material (with a few modifications / notes-to-self) into a private section of this site's database (which I can't make public because it's copyrighted information). From those organized notes I combined all the information from those books and my own experiences to write the articles. The point of that little aside is that I'm going to post all my notes for Drunella lata to answer Konchu's question. These are mostly just lists of direct quotes of copyrighted material, but since I'm blockquoting and attributing it I think I'm ok here. These are private notes so there are probably lots of typos, the result of my typing them into my database at 110 words per minute while not looking at the screen. Caucci, Al and Nastasi, Bob. 2004. Hatches II. The Lyons Press. (refers to lata itself) Arbona, Fred Jr. 1989. Mayflies, the Angler, and the Trout. Nick Lyons Books. (described for cornuta) Leonard, Justin W. and Fannie A. Leonard. 1962. Mayflies of Michigan Trout Streams. Cranbrook Institute of Science. Habitat: Nymphs occur widely in gravel riffles in steams of all sizes Knopp, Malcolm and Robert Cormier. 1997. Mayflies: An Angler's Study of Trout Water Ephemeroptera . The Lyons Press. Emergence: First week May through second week August Fauceglia, Ted. 2005. Mayflies . Stackpole Books. (discusses the lata synonyms together with walkeri) Swisher, Doug and Carl Richards. 2000 edition. Selective Trout. The Lyons Press. Emergence: July 1 to August 10 | |
Jason Neuswanger, Ph.D. Troutnut and salmonid ecologist | ||
Konchu | March 8th, 2007, 9:24 pm | |
Site Editor IndianaPosts: 505 | Troutnut: you ARE nuts! (We're not worthy, we're not worthy) Where were you when I was trying to take notes in college? | |
Troutnut | March 9th, 2007, 5:23 am | |
Administrator Bellevue, WAPosts: 2737 | Probably taking notes in college. :) | |
Jason Neuswanger, Ph.D. Troutnut and salmonid ecologist | ||
Martinlf | March 9th, 2007, 6:30 am | |
Moderator Palmyra PAPosts: 3233 | Wow. This is amazing. And I didn't notice any typos in my quick scan through. | |
"He spread them a yard and a half. 'And every one that got away is this big.'" --Fred Chappell | ||
Troutnut | March 9th, 2007, 8:03 am | |
Administrator Bellevue, WAPosts: 2737 | Must have been a very quick scan! | |
Jason Neuswanger, Ph.D. Troutnut and salmonid ecologist | ||
Konchu | March 9th, 2007, 9:37 am | |
Site Editor IndianaPosts: 505 | FROM CAUCCI & NASTASI (2004): Finally, we believe that cornuta's obscurity was due to a simple case of mistaken identity. We are convinced that even the most serious experts have mistaken the activity of cornuta for that of Ephemerella attenuata. This got my attention. What does everyone think about this speculation? Has anyone followed up on this? It seems possible to me... | |
Taxon | March 9th, 2007, 11:28 am | |
Site Editor Royse City, TXPosts: 1350 | FROM CAUCCI & NASTASI (2004): Konchu- It certainly sounds plausible, but I believe the author of that passage was talking about mistaking the emerging duns of Drunella cornuta for those of Attenella attenuata, rather than mistaking identification of immatures of the respective species. Further, I believe that “the most serious experts” wasn’t necessary a reference to professional taxonomists specializing in mayfly classification. In any event, the full pertinent passage reads as follows: Finally, we believe that cornuta’s obscurity was also due to a simple case of mistaken identity. We are convinced that even the most serious experts have mistaken the activity of cornuta for that of Ephemerella attemuata. Both are typical of the Blue-Winged Olive varieties that hatch in the morning hours during the season. To get to the base of the angler’s identification problem, let’s scrutinize these similarities. To the untrained eye, cornuta and attenuata look alike. Yet, up close, they are physically different and they have different emergence traits. The cornuta dun is larger, averaging about 9 mm, while attenuata averages around 7 mm (a difference of one hook size). Although the wings are identical in color, the olive body hues are each different. | |
Best regards, Roger Rohrbeck www.FlyfishingEntomology.com | ||
Konchu | March 9th, 2007, 2:18 pm | |
Site Editor IndianaPosts: 505 | absolutely it wouldn't be the nymphs lata is relatively more common, so their statement seems to hold true; i was curious about historic/present confusion of hatches of the two regarding who's an expert, that's usually someone at least 50 miles away from home... ;) PS, Thanks, Taxon, for the complete quote. | |
GONZO | March 17th, 2007, 3:10 pm | |
Site Editor "Bear Swamp," PAPosts: 1681 | Konchu and Roger- I noticed this thread last week during a break from skiing, but I didn't have time to reply. The former D. cornuta is a personal favorite of mine, and I have more than twenty years of notes that reflect my pursuit of this hatch in the waters of PA and NY. Historically, the new D. lata synonyms make for a fascinating study and are the source of much confusion in the fly-fishing (and scientific) literature. As for the supposed D. cornuta/A. attenuata confusion, you are right that this probably pertained only to adults. (The nymphs would be pretty hard to mistake for one another.) Early fly-fishing entomologies like Schwiebert's Matching the Hatch and Flick's Streamside Guide (2nd edition) refer to attenuata, but make no mention of Drunella (then Ephemerella) species like cornuta, longicornus, cornutella, or lata. Hence, a casual encounter with any of these morning-emerging olives might easily have been attributed to attenuata. The original version of Swisher and Richard's Selective Trout (1971) describes the Midwestern activity of lata, but continues to perpetuate the myth of the significance of attenuata by describing it as an Eastern "superhatch" (with no mention of cornuta). Schwiebert's later volume Nymphs (1973) includes A. attenuata as well as D. cornuta, D. longicornus, and D. lata (at the time, all were under the genus Ephemerella). Curiously, Schwiebert attributes the "extremely important hatch in June" on the Brodheads to lata. This was mistaken identity at the time (it would have been cornuta under the former taxonomic divisions), but now is strangely correct. The original Caucci and Nastasi Hatches (1975) was the first angling entomology to properly attribute and describe the cornuta activity in the East. While they discounted the presence of lata in the East, this morphotype is also found in many Eastern waters, although its prominence usually doesn't approach what is typically accorded to it in the Midwest. (And while the authors mention attenuata as a species of local importance, I'm reasonably sure that their photograph of an "attenuata" dun is really that of a Drunella dun.) Around that time, cornuta came to be recognized as the most important morphotype in the East, and lata was considered the most important type in the Midwest (though both of the former species were recorded by scientists in both regions). Upon reviewing the citations that Jason has included above, I find the Caucci/Nastasi and the Arbona information to be the most accurate based on my experience and observations. Later books (like Knopp's or Fauceglia's) mostly parrot this information. Thomas Ames' recent Hatch Guide for New England Streams, which is otherwise excellent, describes the Eastern Drunella activity as primarily a midsummer hatch of small (#16-18) flies occurring around noon. This is a very limited assessment and doesn't account for the earlier hatch of larger and (usually) more important flies (the former cornuta). I can only assume that his experience is limited to the later lata hatch in cold tailwaters (hence, the atypical emergence timing). As mature nymphs, and under magnification, their frontal horn differences are quite distinctive. Roger, while these differences in horn (frontoclypeal projection) length were often used as a character to differentiate between the former species, Jacobus and McCafferty (Revisionary Contributions to Drunella [Ephemeroptera], J.N.Y.E.S. 112[2-3]: 127-147, 2004) say that this is a variable trait: "Based on specimens we have examined...the frontoclypeal projection characterizations attributed to D. lata, D. cornutella, D. cornuta, and D. longicornus appear arbitrary." They do say, however, that "Frontoclypeal projection development is usually consistent within samples taken from a single population...." Other morphological characters that have been used as a means of differention are also revised as follows: Epicranial sutures--consistent across morphotypes of former species. Median ocellar spine--relatively well-developed, but variable in terms of bluntness or sharpness. Third genital forceps segment (adult males)--generally about three times or more as long as wide, but variable within that range among populations and individuals. In addition, Koss (1968) and Studemann and Landolt (1997) provided equivalent egg data for D. cornuta, D. cornutella, and D. lata. And equivalent, highly variable adults were reared from each larval morphotype. The best attempt I can make at answering Konchu's original question is to relate the information from my own notes, which reflect the sequential activity of three of the synonyms as found in my favorite streams--the former D. cornuta, D. cornutella, and D. lata. Keep in mind that this information is specific/anecdotal to the Poconos and Catskills (though similar activity occurs in much of the East), and that the former species are distinguished here (for my purposes) primarily by size, color, and emergence period. Here's how the annual activity shakes out in a favorite Pocono watershed: Olive Morning Dun (formerly D. cornuta). This is the main event and the heaviest Drunella hatch of the season. The large (9-11mm) nymphs are somewhat concolorous, ranging from tannish olive to dark olive-brown. Some display brownish banding on the legs and others do not. The smaller male nymphs are somewhat darker, have large eyes, and the developing genital forceps are visible on mature specimens. Males seem to dominate the early part of the hatching period. On lower mainstem waters, the hatch usually commences in late May--earliest date on 5/21 and latest on 6/9. Because this same hatch extends well into the small cold headwaters, hatching there can be as late as early July (latest headwater date, 7/11). Under ideal conditions, the peak hatching occurs at a water temperature of around 58 degrees F. When the weather is either too cold or too warm to reach this range, the hatch occurs when the water temperature makes the greatest movement toward the ideal. This usually translates into a morning peak somewhere between 8:00-10:00 am. Hot spells concentrate hatching closer to daybreak, and unusually cold weather may cause sporadic hatching well into the afternoon. (The latest daily peak I have experienced was around 2:00 in the afternoon on a very cold and rainy day.) In a given stretch, the hatch lasts for 4-7 days, with females becoming more prominent during the latter part of the hatching period. In the Poconos, this is one of the most reliable, predictable, and productive mayfly hatches. Little Olive Morning Dun (formerly D. cornutella). This smaller hatch (in both size and concentration) follows the former cornuta, and in all respects is a miniature version of that activity. The nymphs are virtually identical in appearance except for their size (6-8mm). Due to warmer weather, the hatch usually occurs earlier in the morning and is typically finished in the lower mainstem water by sometime in mid-July. (The latest date I have found the nymphs in the mainstem is 7/20.) Because of their extreme similarity, I was not surprised to find that they are considered to be a diminished component of the earlier hatch. Interestingly, I have not found this hatch to follow immediately on the heels of the earlier hatch. Instead, there seems to be a lag of a few weeks before the smaller synonym gets started. Dark Olive Morning Dun (formerly D. lata). This is the most curious and puzzling of the three components of the new lata hatch. In size, it usually matches the former cornutella at maturity (6-8mm), but its coloration (in Pocono waters) is very different. Descriptions and photographs of the former lata larval morphotype often have contrasting bright or light colors on the pronotum, femurs, and sometimes bright spots or streaks near the rear of the abdominal tergites. The contrasting colors can be dark brown with orange or crimson accents, or very dark olive (almost black) with bright green accents. In the Pocono streams, the latter color combination predominates. Because hatching typically occurs in July heat, the daily timing is often close to daybreak. This is the smallest and least significant of these synonymous hatches. It usually peaks in mid- to late July, and the latest that I have found these nymphs in the mainstem is 7/26 (although they may extend into August in some years). While all of the Drunella duns are a fairly light green immediately upon emergence, this one is typically darker and ripens into a very dark olive/grey dun in a short time. In summary, I find these Drunella olives to be very important in many (if not most) Eastern trout streams. They are found in both acid and alkaline waters (the acidic Tobyhanna Creek and the alkaline Penn's Creek are two contrasting examples with good populations). They thrive in large mainstem waters as well as in tiny tributaries. Only streams without fastwater stretches seem to lack populations. (I have never found a good population in flat, weedy limestoners, while limestone streams with extensive fastwater stretches--like Penn's Creek--usually have good populations.) I have fished them in streams as diverse as the Delaware River, the Beaverkill, the Esopus, the Brodheads, the Tobyhanna, the Lehigh River, the Yellow Breeches, Mountain Creek, Penn's Creek, Fishing Creek (both the one in Columbia County and the more famous one in Centre/Clinton County), and even in Logan Branch and upper Spring Creek. They are a favorite hatch because the trout love them, and they have been very rewarding to study. The main reason that they are not better known or understood is that the heaviest hatches occur on mornings when most fly-fishers are pursuing the famous evening hatches--Sulphurs, Yellow Quills, March Browns, Light Cahills, Slate Drakes, Green Drakes, and Pale Evening Duns. This period (late May through early June) constitutes "prime time" for mayfly hatches on many Eastern waters, but the morning hours are often neglected by dedicated hatch-matchers. This situation allows the former D. cornuta to hide in plain sight. Personally, I'm quite content with its relative obscurity. Gonzo | |
Konchu | March 30th, 2007, 6:58 pm | |
Site Editor IndianaPosts: 505 | Gonzo: Thanks for the treasure trove of information. I hope to spend more time with it in a few days, in order to sort some things out. I think your encyclopaedic dedication may match that of Troutnut. -- konchu | |
GONZO | March 31st, 2007, 8:20 am | |
Site Editor "Bear Swamp," PAPosts: 1681 | Thank you, Konchu. But Jason is a true "phenom." It's scary to think what he'll become in ten or twenty years. The fish will have to be warned! :) | |
DarkDun | May 2nd, 2007, 5:36 pm | |
Posts: 16 | I never realized my original post would draw so much attention, but this is a serious hatch in every part of the US. Lots of new info for me such as the Cornuta hatches of PA which I have fished several times and assumed them to be Lata. Also the Longicornus of NC, a new insight for me into our local NC hatches. | |
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